Greek police started removing migrants from the Greek-Macedonian border on Tuesday after additional passage restrictions imposed by Macedonian authorities left hundreds of them stranded, sources said.

The migrants had squatted on rail lines on Monday after attempting to push through the border to Macedonia, angry at delays and additional restrictions in crossing.

Greek police and empty buses had entered the area before dawn, a witness said. In one area seen from the Macedonian side of the border, about 600 people had been surrounded by Greek police, the witness said.